Associate Professor Jessica Rettig, Chair
Professors Eric C. Liebl, Tom D. Schultz ; Associate Professors Warren D. Hauk, Rebecca N. Homan, Kristina S. Mead, Jessica E. Rettig, Laura A. Romano, Geoffrey R. Smith, Jeffrey S. Thompson, Christine L. Weingart, Lina I. Yoo; Assistant Professors Ayana Hinton, Clare C. Jen, Andrew C. McCall, Heather J. Rhodes; Visiting Assistant Professor Tessa L. Carrel; Academic Administrative Assistant Jenny Etz; Lab Manager/Bioreserve Manager Whitney Stocker
The Department of Biology endeavors to provide a comprehensive foundation in concepts and skills across the breadth of biology through an introductory core of three courses that prepares students for a deep exploration of sub-disciplines and research methods through subsequent advanced courses. The core covers the major concepts of biology and basic skills of acquiring and processing information, problem solving, and analyzing data. Our program then allows students the flexibility to explore specific areas of biology in depth through a suite of advanced courses in which they can expand and apply their knowledge and skills.
The major prepares students for careers in science and related fields as well as graduate and professional schools (including pre-medical, pre-dental and pre-veterinary studies), while allowing students the flexibility to design the program that best suits their specific interests and career goals. In addition, biology majors are offered the opportunity to collaborate with faculty in research and laboratory instruction; to present exceptional work at professional meetings; and to assist in the maintenance of the 350-acre Biological Reserve. Related programs in Medicine, Dentistry, Medical Technology, Forestry and Natural Resources are described under Pre-Professional Programs.
Requirements for Biology Majors. Students majoring in Biology (B.A. or B.S.) should complete the three Biology core courses, Introduction to the Science of Biology (150), Cell and Molecular Biology (201) and Ecology and Evolution (202) preferably by the end of their second year. The Department of Biology recommends strongly that students earn a C or better in each of the three core courses before proceeding to 300-level courses. The major requires two semesters of Principles of Chemistry (CHEM 131 and 132; grades of C or better are recommended strongly), five or six 300-level advanced courses (depending on the degree sought), one of which must be designated a "biological diversity" course. Advanced Senior Research (452) is credited as a 300-level course. Any combination of advanced courses may be taken to provide a concentrated preparation in a specific discipline or a broader survey of advanced topics. Directed Study (361, 362), Independent Study (363, 364), and Senior Research (451) are not counted as 300-level advanced courses toward the requirements for majors. It is strongly recommended that majors consult with a Biology advisor in order to design the most appropriate suite of advanced courses for that student.
Student majoring in Biology must satisfactorily complete Biology Assessment I (BIOL 300 - core curriculum assessment exam) and Biology Assessment II (BIOL 301 - senior interview) in order to graduate.
Bachelor of Arts in Biology. The requirements for the Bachelor of Arts degree in Biology consist of the three core courses (Biology 150, 201, 202), five 300-level biology courses (one of which must be a designated "biological diversity" course), and one year of Principles of Chemistry (Chemistry 131 and 132). Advanced Senior Research (452) is credited as a 300-level course. Directed Study (361, 362), Independent Study (363, 364) and Senior Research (451) are not counted as 300-level advanced courses.
Bachelor of Science in Biology. The requirements for the Bachelor of Science degree in Biology consist of the three core courses (Biology 150, 201, 202), six 300-level biology courses (one of which must be designated a "biological diversity" course), one year of Principles of Chemistry (Chemistry 131 and 132) and five additional courses in mathematics or science. Advanced Senior Research (452) is credited as a 300-level course. Directed Study (361, 362), Independent Study (363, 364) and Senior Research (451) are not counted as 300-level advanced courses. Students pursuing a B.S. in biology increase their experience in biology and become more broadly trained in the sciences. Thus these students fulfill the B.S. by taking an additional biology course beyond the requirement for a B.A. and by taking five additional non-biology science or math courses, with no more than three taken in a single department or program. Because the B.S. trains students more liberally as scientists, students are encouraged to fulfill these non-biology courses by selecting classes that "do" science, such as classes that include laboratories. However, any non-biology course within the science division, any environmental studies (ENVS) science course, or Applied Anatomy (PHED 204) will fulfill the B.S. requirement. Biology majors preparing for medical school or most graduate programs, and enrolling in Fall 2012, are advised to take Intermediate Organic Chemistry (CHEM 251) and Biochemistry (CHEM 258), General Physics (Physics 121-122), and two semesters of college-level math (e.g., Calculus (Math 121,123,or 124) or Statistics (Math 102)).
The Biology Minor consists of six courses in Biology. All students electing a minor in Biology must complete the three core courses (Biology 150, 201, 202) plus one semester of chemistry (Chem 131) and three additional 300-level courses. Senior Research (451), Advanced Senior Research (451), Directed Study (361, 362) and Independent Study (363, 364) are not counted as 300-level advanced courses. One of the 300-level courses must be designated a "biological diversity" course.
Biological Diversity Courses: Courses that fulfill the biological diversity requirement emphasize the importance of scientific studies at the level of the whole organism. In these courses students gain a holistic perspective on the study of organisms, explore a variety of living forms through a broad survey of taxa, and evaluate the role of phylogenetic history in taxonomy. Students also use careful observation to learn morphology and diagnostic traits, identify organisms to meaningful taxonomic units, and learn the principles of scientific nomenclature. The 300-level biology courses designated as biological diversity courses are: BIOL-312 Herpetology, BIOL-313 Vertebrate Zoology, BIOL-317 Diversity of Microorganisms, BIOL-320 Plant Systematics, BIOL-326 Plant Evolution & Reproduction, and BIOL-327 Biology of Insects.
Students may complement their major in biology through study off-campus. Denison University is a member of several consortia that offer course credit through off-campus programs. Those with course offerings relevant to Biology students include: the School for Field Studies, the SEA Semester, the Organization of Tropical Studies, the Duke University Marine Laboratory, the Semester in Environmental Science, the Oak Ridge National Laboratory, the Institute for Study Abroad, Denmark's International Study Program, and the Associated Colleges of the Midwest Wilderness Field Station. The Department of Biology is committed to awarding credit for courses offered through those programs that provide a sufficient focus on biological concepts and methods (lecture and laboratory). With prior approval from the department, a maximum of two off-campus courses may be credited as advanced electives and counted toward the requirements of the major. The Richard C. and Linda G. Seale Scholarship provides support to qualified Denison students for participation in summer courses at the Duke University Marine Laboratory. Financial aid may be available for other off-campus programs.
Courses for Non-Majors. Non-majors are invited to take Modern Topics in Biology (100, 103, or 104), courses designed to explore scientific inquiry and biological concepts through specific topics in the instructor's area of expertise. Introduction to the Science of Biology (150) may be taken by any student to fulfill the General Education requirement for science, but is recommended only for Biology and related majors. Students with Advanced Placement or International Baccalaureate credit in Biology may receive credit for Biology 100 without General Education credit. In addition, students who have had extensive biology training in secondary school may petition the Biology department for Biology 100 credit without General Education credit. However, such petitions must be made before the completion of the student's third year at Denison.
Biology and Environmental Studies. Students with an interest in both Biology and Environmental Studies may pursue a major in Biology with a minor in Environmental Studies, or a major in Environmental Studies with a concentration in biology. Students are advised to choose the programmatic path that best suits their post-graduate goals, and to seek early consultation with faculty in Biology and/or Environmental Studies.
Biology and Neuroscience. Students with an interest in both Biology and Neuroscience may pursue a major in Biology with a concentration in Neuroscience. Students interested in this option should consult with a faculty member early in their career.
Modern Topics in Biology (BIOL-100). This course for non-majors is intended to promote scientific literacy. Topics will vary with the instructor, but each edition of the course will focus on a specific topic as a vehicle for exploring the essentials of biology and the scientific method. This course satisfies the G.E. lab science requirement. Biology 100 may not be counted toward the major in biology. Three class periods and one laboratory weekly. 4
Modern Topics in Biology (BIOL-103). This course for non-majors is intended to promote scientific literacy and quantitative reasoning. Topics vary with the instructor, but each edition of the course will focus on a specific topic as a vehicle for exploring the essentials of biology and the scientific method. This course satisfies the G.E. lab science requirement as well as the quantitative reasoning requirement. Biology 103 may not be counted toward the major in biology. Three class periods and one laboratory weekly. 4
Modern Topics in Biology (BIOL-104). This course for non-majors is intended to promote scientific literacy and oral communication. Topics will vary with the instructor, but each edition of the course will focus on a specific topic as a vehicle for exploring the essentials of biology and the scientific method. This course satisfies the General Education lab science requirement as well as the oral communication requirement. Biology 104 may not be counted toward the major in biology. Class meets for two (80 minute) or three (50 minute) periods per week plus a three-hour laboratory. 4
Introduction to the Science of Biology (BIOL-150). This course is the first biology course biology majors take and is not recommended for non-majors. It is a course that introduces students to core concepts of modern biology through active participation in biological investigations. Topics include reproduction, Darwinian evolution, energetics, organisms' response to stimuli, and organismal structure and function. Imbedded throughout the course are many of the skills expected of practicing biologists including the ability to develop hypotheses and analyze and interpret data, the ability to present scientific data, scientific writing, and a familiarity with the scientific literature. This course satisfies the Quantitative Reasoning requirement. Three class periods and one laboratory weekly. Offered Fall and Spring semesters. 4
Cell and Molecular Biology (BIOL-201). The study of cellular structure and function from a molecular perspective. The organization and molecular composition of cells is examined, with a particular emphasis on distinctions between the prokaryotic and eukaryotic domains. Major cell functions studied include membrane transport, signal transduction, and eukaryotic cell cycle regulation. The molecular basis of genetic expression is addressed, including topics such as DNA replication, transcription, translation, and associated regulatory processes. Evolution is also explored from the perspective of cellular and molecular biology. Prerequisites: BIOL 150 or Consent of Instructor. Chemistry 131 pre- or co-requisite. Three class periods and one laboratory weekly. Offered Fall and Spring semesters. 4
Ecology and Evolution (BIOL-202). This course explores the fundamental biological concepts of ecology and evolution and integrates them in a study of the interactions between organisms and their environment and how those interactions shape the history of life on Earth. With a thorough understanding of population genetics and natural selection, this course addresses ecological questions at the level of the individual, population, community and ecosystem. A common thread that binds the course is the role of deterministic and stochastic processes in shaping ecological systems and macroevolutionary patterns. Prerequisite: BIOL 150 or Consent of Instructor. Three class periods and one laboratory weekly. Offered Fall and Spring semesters. 4
Minor Problems (BIOL-250). A research problem (library or laboratory) of limited scope which provides the opportunity for the qualified student to extend his or her interest beyond the limits of particular course offerings. Does not count toward minimal department requirements. 1 or 2
Biology Assessment I (BIOL-300). A pass/fail course used to track all biology majors' completion of the required assessment exam covering the Biology core. Earning the required S (pass) in this course entails attending an information session explaining the exam and taking the assessment exam in good faith. Offered Fall and Spring semesters. 0
Biology Assessment II (BIOL-301). A pass/fail course used to track all biology major's completion of the required senior interview. Earning the required S (pass) in this course entails attending an information session explaining the biology department's senior interview and completing the senior interview in good faith. Seniors enroll in BIOL 301 in their last semester at Denison. 0
Biochemistry (BIOL-302). A study of the chemical and physiochemical properties of living organisms. Concepts will be developed through a study of the physical and chemical properties of biological compounds and integration of various metabolic pathways in an attempt to understand the dynamics of living systems. The laboratory will include the isolation and study of properties of biological compounds. Prerequisites: CHEM 224 and Biology 201. The department strongly recommends that students enrolling in this course have earned a grade of C or higher in Organic Chemistry II (CHEM 224) and Molecular Biology (BIOL 201). Offered in the fall semester (also as Chemistry 302). Three class periods weekly plus laboratory. Safety glasses required. Note that due to curricular changes, BIOL 302 will not be offered after 2012-2013. 4
Biodiversity Through Time (BIOL-308). An introduction to the study of fossil invertebrates with emphasis on preservation, taphonomy, diversity trajectories through geologic time, evolutionary mechanisms, extinction, paleobiology and paleoecology. Special emphasis will be placed on using fossils to interpret ancient depositional environments. Labs will introduce the student to the major invertebrate phyla commonly preserved in the geologic rock record. Prerequisite: GEOS 210 or BIOL 202. (Normally offered Fall Semester in alternate years) 4
Computational Biology (BIOL-309). As large and complex data sets have become more prevalent in biology, computer algorithms for analyzing the data have become critical, driving the need for scientists with expertise in both fields. This interdisciplinary course will explore this intersection, examining the biology and the computational methods behind a variety of interesting and important problems. Students will initially work with a single instructor to build a background outside of their own discipline (Biology students with a Computer Science instructor, CS students with a Biology instructor), followed by a merging of the two groups into a single team-taught class, which will investigate a series of biological problems with a computational focus. The laboratory portion of the course will involve students working together in multidisciplinary groups to design algorithms to investigate these problems, as well as undertaking a self-designed “capstone” project at the end of the term. BIOL 309 Prerequisites: Biology core or consent. Students are also strongly encouraged to have taken CS 111. 4
Wetland Ecology (BIOL-310). This course is a comprehensive study of wetland ecology, management, and policy. The main emphasis is on biological, chemical, and physical aspects of major wetland ecosystems found in North America. The course also deals with valuation, classification, and delineation of wetlands. A significant portion of the course focuses on local and regional wetland ecosystems: their history, ecology, and current status. Labs will be field-based explorations of the biology, chemistry, and ecology of these regional wetlands. Prerequisite: BIOL 202 or consent. 4
Herpetology (BIOL-312). Herpetology is the study of amphibians and reptiles, two diverse taxonomic groups that share the characteristic of being ectothermic vertebrates. This course will examine three main areas of herpetology: 1) the evolutionary relationships and biogeographical histories of these taxonomic groups, 2) comparative physiology, and adaptations of amphibians and reptiles to their natural environments, and 3) the ecology of the herpetofauna, as well as conservation issues, with a focus on amphibians. Emphasis will be placed on the critical reading of primary literature on both historical and current issues in herpetology, as well as on gaining hands-on experiences with amphibians and reptiles. Laboratories will include comparative studies of physiology and field studies of native Ohio amphibians and reptiles, making extensive use of the Denison University Biological Reserve. Prerequisites: Biology core or consent. Herpetology qualifies as a "biological diversity" course for the major. 4
Vertebrate Zoology (BIOL-313). In this course we investigate the biology of vertebrates. In particular, we will be considering the many ways in which vertebrates interact with and respond to their environment, and thus this course will emphasize the evolution, ecology, and physiology of vertebrates. Laboratories will focus on the biology of local vertebrates, and will consist of field and laboratory exercises, as well as field research projects. We will make extensive use of the Denison University Biological Reserve. Prerequisite: Biology core or consent. Vertebrate Zoology qualifies as a "biological diversity" course for the major. This course satisfies the Oral Communication requirement. 4
General Microbiology (BIOL-315). This is an introductory course in microbiology emphasizing the general structure, occurrence, habitats, and types of bacteria, viruses, and eukaryotic microbes. Mechanisms of pathogenicity and host defense strategies also are discussed. The course structure includes small group activities, student presentations, traditional lectures, and discussions of scientific literature. Laboratory emphasis is placed on the fundamental techniques of microbiology (i.e., staining, microscopy, and streak plating) and self-designed investigative labs. Students may either take General Microbiology (BIOL 315) or Diversity of Microorganisms (BIOL 317) during their academic career, but not both courses. Prerequisite: Biology core or consent. 4
Virology (BIOL-316). Virology is a course that will examine the diversity of plant, animal, and bacterial viruses. Emphasis will be placed on topics such as molecular interactions between the host and virus, the genetics and chemical nature of viruses, and the replication strategies of viruses. This course also will examine how viruses cause disease, how they are used in biotechnology, and their overall impact on society. The structure of the course will provide peer learning activities, class discussions of primary literature, and traditional lectures. The structure of the laboratory will allow students to develop and test their own hypotheses while learning bacteriophage and tissue culture techniques. Prerequisite: Biology core or consent. 4
Diversity of Microorganisms (BIOL-317). This course examines the remarkable environmental, physiological, and metabolic diversity of prokaryotic and eukaryotic microorganisms (i.e., bacteria, protists, algae, & fungi). More specifically, diversity will be studied in terms of taxonomy and phylogeny, the ability of species to live in various environments, and the application and value of genomics in diversity. Emphasis will be placed on the reading of primary literature, and on using that information to make connections with class lectures and generate hypotheses that will be tested in the laboratory. The structure of the course includes traditional lectures, class activities, and student presentations. Prerequisite: BIOL 150, 201, & 202, or consent. Diversity of Microorganisms qualifies as a "biological diversity" course for the major and minor. Students may either take General Microbiology (BIOL 315) or Diversity of Microorganisms (BIOL 317) during their academic career, but not both courses. 4
Plant Systematics (BIOL-320). In Plant Systematics students learn how major groups of vascular plants are classified, named, and identified. We study approximately 50 plant families concentrating on native representatives (using living plant material whenever possible), learn how to use keys and floras to identify local species, and learn how to find information about plants in traditional and electronic sources. Understanding evolutionary relationships among the families studied is a central theme. This course provides important background for students planning to do fieldwork in ecology, plant-animal interactions, environmental education, and related subjects. Prerequisite: Biology core or consent. Plant Systematics qualifies as "biological diversity" course for the major. 4
Plant Ecology (BIOL-321). In this course we will explore how plants interact with their environments and with other organisms, including man. We will begin at the individual level, learning how plants obtain resources from abiotic sources and through mutualistic interactions with bacteria and fungi. We will also consider how the theories of plant community ecology developed in the early 20th century and why they are pertinent today. Students will also have the opportunity to read and critique primary literature from leading journals in the field. Finally, we will develop several projects to be completed at the Denison Biological Reserve during the term for lab projects. These projects will be student-inspired and driven, with the hopes that they will contribute to our understanding of our immediate surroundings at Denison. Prerequisite: Biology core or consent. 4
Developmental Biology (BIOL-324). Every multicellular organism begins its life as a single cell. Developmental biology is the study of the progression from this single cell to a complex, multicellular organism. Recently the powerful tools of molecular biology have linked the fields of embryology and genetics to reveal how cells, tissues, organs, and organisms develop. Especially striking is the conservation of molecules and mechanisms that underlie developmental processes in different organisms. This course provides an overview of the major features of early embryonic development in animals, and the mechanisms (molecular mechanism when known) that underlie them. We focus on two major aspects of developmental biology: (1) How is the basic body plan established? How does the basic organization of the embryo arise from the fertilized egg? What are the cellular mechanisms underlying morphogenesis and the appearance of pattered structures in the embryo? (2) How do parts become different in the embryo? Prerequisite: Biology core or consent. 4
Genetics (BIOL-325). This course provides a detailed and up-to-date understanding of genetics, an appreciation of how genetics affects our lives everyday from the supermarket to the doctor's office, and a realization of the applications of genetics to virtually every discipline of biology. We focus on three major areas of genetics: (1) Molecular genetics: Thinking about genetics on the DNA level - everything from DNA sequencing to mutagen testing. (2) Mendelian genetics: Thinking about genetics on the gene level-everything from inheritance to recombinational mapping. (3) The application of both molecular and Mendelian genetics to study biological processes. We start by seeing how genetic techniques can be used to dissect almost any biological process and end up answering questions such as: How does genetic disease screening work? How are genes cloned from complex organisms such as mice or even humans? How does gene therapy work? In the laboratory we carry out both molecular experiments and classical genetic experiments. Prerequisite: Biology core or consent. 4
Plant Evolution and Reproduction (BIOL-326). In this course we will explore the evolutionary relationships and histories among the major groups of plants, both terrestrial and aquatic. We will pay particular attention to their modes of reproduction and the structures that facilitate gamete production and dispersal. We will learn how plant physiology and developmental mechanisms have allowed taxa to persist or make major transitions among different environments over time. Class reading material will consist of the primary literature and will be presented by students every week. For the laboratory component we will have one overnight trip to Hocking Hills on a weekend in September to examine and identify plants in their natural habitat, as well as shorter trips to Blackhand Gorge and the Dawes Arboretum. We will also plan together and complete a semester-long project on the effects of environment on the development of reproductive structures in the model plant, Arabidopsis thaliana. Plant Evolution and Reproduction qualifies as a "biological diversity" course for the major. Prerequisite: Biology core or consent. 4
Biology of Insects (BIOL-327). In this course we will explore the world of insects and their interactions with other species. Our central focus will be to survey insect diversity and explore how various orders, families, and species are adapted through evolution to their specific environment. But we will also use that diversity as a lens through which we will examine major concepts in biology. Topics of discussion will include the following: plant-insect coevolution, mating systems, anti-predator defenses, eusocial behavior, parasitism, disease transmittance, insect conservation, and control of agricultural pests. Laboratory will involve collecting insects in the field (including at times outside of class hours), identification, and preparing a collection. Biology of Insects qualifies as a "biological diversity" course for the major. Prerequisite: Biology core or consent. 4
Evolutionary Developmental Biology (BIOL-333). Evolutionary developmental biology (or "evo-devo") is an exciting interdisciplinary field of research that seeks to understand how developmental mechanisms have evolved to produce differences in the anatomy, physiology, and behavior of organisms. This course will begin with an overview of basic concepts in developmental biology. Students will then learn about the genes responsible for specific processes and examine the functional consequence of changes in their expression during embryonic development. (For example, students will learn about the genes that regulate eye development in vertebrates, and then examine how changes in their expression have led to organisms with different types of eyes, or no eyes at all!) In lab, students will conduct a semester-long project designed to provide insight into the process by which biologists explore the evolution of developmental mechanisms. In particular, the project will involve cloning genes and analyzing their DNA sequences using a variety of bioinformatic tools. Prerequisite: Biology core or consent. 4
Comparative Animal Physiology (BIOL-334). This course is a comparative study of how animals perform their life-sustaining functions. We'll use a wide variety of animal examples to explore the physiology of metabolism, digestion, thermoregulation, muscles, and the cardiovascular, respiratory, and osmoregulatory systems. This course will examine the adaptive significance of physiological traits at the molecular, tissue, organ and whole organism level. In addition, it will stress the ways that physiology and ecology interact, currently and over evolutionary time. Students will participate in several course labs and then design their own physiology experiments. Students may not dual enroll in BIOL 334 and BIOL 335 in the same semester. Prerequisite: Biology core or consent. 4
Human Physiology (BIOL-335). In this course we will examine the physiology and anatomy of the major systems of the human body, including the cardiovascular, nervous, muscular, endocrine, renal, and digestive systems. We will study how the body functions to sustain life and maintain homeostasis from the level of single cells up to multi- organ systems. The course will also incorporate discussions of disease processes when the body fails to function as it should. Students will participate in lab exercises examining the function of their own human bodies and will design their own physiology experiments. Students may not dual enroll in BIOL 334 and BIOL 335 in the same semester. Prerequisite: Biology core or consent. 4
Animal Behavior (BIOL-340). In this course we study the proximate and ultimate causes of animal behavior from an evolutionary perspective. Topics include the genetic, developmental and neural bases of behavior as well as behavioral strategies of habitat choice, foraging, defense, courtship, parental care and sociality. The laboratory will include several multi-week experiments designed to test hypotheses concerning behaviors observed in the field and lab. There will be a strong emphasis on data analysis and interpretation, and use of the primary literature. Prerequisite: Biology core or consent. 4
Immunology (BIOL-341). This course is a study of concepts in immunology, focusing on the cellular and molecular aspects of the immune system in humans and other animal models. We will delve into subjects allowing students to understand the fascinating and complex mechanisms with which our immune systems defend our bodies against a constant barrage of infectious microorganisms. Topics covered include immune cell development and function, specific and non-specific immune responses to infection, immunogenetics, vaccination, and clinical disorders of the immune system such as allergies, immunodeficiency diseases, and autoimmunity.Laboratory exercises will utilize immunological techniques to address questions pertaining to the molecular function and specificity of the immune system. Prerequisite: Biology core or consent. 4
Eukaryotic Cell Biology (BIOL-345). This course will be an in-depth examination of fundamental cellular functions, with an emphasis on how disturbances in these functions lead to disease. Areas covered in the course include intracellular trafficking, cytoskeleton and cell motility, adhesion, signal transduction, cell cycle, and apoptosis. Laboratories will involve learning current methods to analyze biological processes in cells. Prerequisite: Biology core or consent. 4
Introduction to Neurophysiology (BIOL-349). We will use neurophysiology and neuroanatomy to understand the links between molecules, cells, systems, and ultimately behavior. The course will start with an exploration of neurons and signaling within and among cells. We will then examine some sensory and motor systems. The last portion of the course will examine the whole animal in a neurophysiological context. The classroom portion of the course consists of lectures, discussion of the text and of research articles, problem sets, analysis of case studies, and other activities. The laboratory component will involve a mixture of behavioral, anatomical, and physiological studies on vertebrate and invertebrate animals, electronic modeling of nerve circuits, and computer simulations of nerve activity. The labs are designed to introduce students to some fundamental neurophysiological techniques and to a variety of study organisms, and to strengthen experimental design and analysis skills. Prerequisite: Biology core or consent. 4
Genomics (BIOL-350). Genomics is the study of genomes, the entire collection of genetic information found in a specific organism. This field of study attempts to understand how all of the genes in a given genome cooperatively function to orchestrate the biological activities within the organism. The genomic DNA sequences of hundreds of species have been determined, including humans, providing a wealth of information about the genetic composition and evolutionary relatedness of species. This course will introduce students to the fundamental concepts in genomics, including how genome sequences are assembled, how potential genes within the genome are identified and characterized, how genomes are organized and regulated, and how genomes evolve. Contemporary papers from the field of genomics will be discussed to complement the concepts addressed in class. The laboratory component of this course will be computer-based, utilizing various online databases and "bioinformatic" programs to carry out a series of projects on genome assembly and compositional analysis. Prerequisites: Biology core or consent. 4
Directed Study (BIOL-361). A research problem (library, field, or laboratory) that provides the opportunity for the qualified student to extend his or her interest beyond the limits of particular course offerings. Does not count toward minimal departmental requirements. 1-4
Directed Study (BIOL-362). A research problem (library, field, or laboratory) that provides the opportunity for the qualified student to extend his or her interest beyond the limits of particular course offerings. Does not count toward minimal departmental requirements. 1-4
Conservation Biology (BIOL-370). Conservation Biology requires the broad use of biological disciplines such as ecology, physiology, genetics, and animal behavior, as well as appreciation of policy issues, to understand and manage biodiversity. In this course, students will learn how to apply these biological tools for the purpose of defining and maintaining biodiversity at many scales. We will also cover human impacts on biodiversity, as well as the link between science and policy in protection efforts. This course will emphasize critical reading of primary literature as well as gaining hands-on experiences with population modeling, and measuring and monitoring local biodiversity. Prerequisites: Biology core or consent. 4
Population and Community Ecology (BIOL-375). In this course, we will examine 1) how populations and communities are structured, 2) how populations and communities change over time, and 3) how populations and communities are influenced by their environment or ecological context. An emphasis will be placed on using primary literature and on doing ecology in the field and lab. Prerequisite: Biology core or consent. This course satisfies the Oral Communication requirement. 4
Evolutionary Biology (BIOL-380). This course builds on BIOL 202 and completes an in-depth survey of evolutionary theory with emphasis on processes that drive organismal change. We examine how molecular technology has impacted the study of evolutionary processes, and how new methods of analysis are changing the study of population genetics, phylogeny construction, adaptive radiation, etc. Experimental design and reading of primary and secondary scientific literature are stressed. Through the course, emphasis is placed on integration of all biological disciplines under the paradigm of evolution. Prerequisite: Biology core. 4
Advanced Biochemistry (BIOL-401). This is a topical course, the content of which will vary from year to year. In general, a detailed look at a variety of recent biochemical topics will be conducted through readings of the primary literature. Laboratory will offer an in-depth, semester-long research experience. Offered Spring semester (also as Chemistry 401). Prerequisite: CHEM/BIOL 302. Safety glasses required. Note that due to curricular changes, BIOL 401 will not be offered after 2012-2013. 4
Senior Research (BIOL-451). For seniors desiring to work on an advanced research problem. Biology 451 is to be taken if no previous work on the specific research project has been accomplished. Students with prior, substantial experience on their research project (such as a summer research experience with a Denison faculty member) may petition to move directly into Biology 452. Prior consent of the advising faculty is required for registration. The grade is determined by the advisor. Completion of Biology 451 does not fulfill an upper-level biology course requirement for the major. 4
Advanced Senior Research (BIOL-452). For seniors working on an advanced research problem. Following the completion of a substantial research experience, such as Biology 451 or a summer research experience with a Denison faculty member, students may take Biology 452. Prior consent of the advising faculty is required for registration. The grade is determined by the advisor. Completion of Biology 452 fulfills one upper- level biology course requirement for the major. Students enrolled in BIOL 452 have the option of purusing senior research with Recognition. Interested students should speak with their research advisor or the Chair of Biology to learn more about the Recognition process and expectations. 4